Discord-Daq «-'e'» mojaq wIrIchtaHvIS lo'meyDaj vIbuS. qen QIj SuStel 'ej meqqu'mo' ghaH, De'wI'wIjDaq mojaqvam QIjbogh ghItlh'e' vIpol. 'ach DaH SuStel latlhpu' joq vIghelnIS.
mu'tlheghDaq «'oH, ghaH, bIH, chaH» lo'lu'chugh SeSorDaq «-'e'» chelnISlu' net Sov. ghantoH: «Ha'DIbaH 'oH vIghro''e'.» qay'be'. ghellu'chugh jaSHa' chelnISlu'. ghantoH: «nuq 'oH vIghro''e'?» qay'be' je.
'ach reH mu'tlheghmeyvetlhDaq De' ngo'Daq ngam mojaqvam, qar'a'? not De' ngo''e' 'oS ghelbogh mu' («'Iv, nuq»), qar'a'? vaj tlhIngan Hol pabHa' bIvlaw' mu'tlheghvam: «Ha'DIbaH 'oH nuq'e'?»
'ach «Hov leng: maQmIgh» (STID) HaStamuchDaq Qoy'lu': «DevwI' ghaH 'Iv'e'?» ‘Who is the leader?’ — vaj latlh lo' ghaj'a' mojaqvam?

Short version in English:
In be-sentences -'e' marks the subject, which SuStel explained as one of the TOPIC uses of the suffix. Topic means that it marks old information, so I concluded logically that question words like nuq and 'Iv cannot be marked by -'e'. But there is a sentence in STID, «DevwI' ghaH 'Iv'e'?» ‘Who is the leader?’ Is the usage of -'e' in be-sentences a different (fourth) usage that is neither topic, focus, nor head-marking in a relative clause? Or is it perhaps focus just in this one example after all? Or can we dismiss it saying that Uhura's Klingon isn't quite perfect and native-like and she made a mistake here (I believe she said this sentence)?

Koran antaŭdankon,
— André